LIVRO Plant Ecology
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LIVRO Plant Ecology


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Second singlet state
First singlet state
Triplet
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t / = 10-3 s2
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Heat
PSII LHC PSI
Fluorescence Fluorescence
Heat
Calvin cycle
HeatHeat
Photochemistry: H2O FdZ P680 P700PQQA QB b/f PC
Chl a*
NADPH
Chl a*
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Fv
Fm
qE
qQ
Fv
Fm'
q
Measuring
beam
Actinic light + periodic saturating light flashes
20 sSaturating
light flash
F0: Fluorescence from the 
antenna
Fm': maximum fluorescence of an
illuminated leaf 
(saturating light flash)
Fv: variable fluorescence during
illumination with actinic light
qQ: (new qp) photochemical 
quenching
qE: (new qNP or qN) non-photo-
chemical quenching at 
great \u2206pH (increase in the
rate constant for thermal
dissipation)
q: total quenching under
continuous light
Fm: maximum fluorescence of a
predarkened leaf
F0
0
A
6
5
4
3
2
1
0
F0
Fm
Fm'
F
F0'
\u2206F
Dark adapted Illuminated
(Fm' \u2013 F0)/Fm = Fv/Fm
(Fm' \u2013 F)/Fm' = \u2206F/Fm'
Optimal quantum efficiency:
Actual quantum efficiency:
qp
qN
NPQ
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B
= (Fm' \u2013 F)/(Fm' \u2013 F0')
= 1 \u2013 (Fm' \u2013 F)/(Fm' \u2013 F0')
= (Fm \u2013 Fm')/Fm'
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