FREE ENTOMOLOGICAL LITERATURE

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FREE ENTOMOLOGICAL LITERATURE 
 
The following lists represent a large amount of duplicate entomological literature in storage in 
Entomology at the National Museum of Natural History of the Smithsonian Institution. This literature is 
available free of charge to researchers visiting the Museum or by mail upon request. We will mail 
reasonable requests (non-excessive amounts) to entomological researchers only, this literature is not to be 
sold. This literature is available on a first come first serve basis. We will mail it as staff time and 
resources allow. 
 
Please send any requests by email to David Furth (furthd@si.edu) 
 
ANOPLURA PUBLICATIONS 
 
Ewing, H. E., 1932. A New Sucking Louse from the Chimpanzee. Proceedings of the Biological Society 
of Washington, 45:117-118. 
 
Ewing, H. E.,1935. The Taxonomy of the Anopluran Genera Polyplax and Eremophthirius, Including the 
Description of New Species. Proceedings of the Biological Society of Washington, 48:201-210. 
 
Johnson, P. T. 1957. The Identity of Pediculus spiculifer Gervais (Anoplura). Bulletin of the Brooklyn 
Entomological Society, 52(1):25-27. 
 
Johnson, P. T. 1957. Description of Two New Species of Eulinognathus Cummings, and Redescription 
of Eulinognathus aculeatus (Neumann) (Anoplura). The Journal of the Egyptian Public Health 
Association, 32(6):273-283. 
 
Scanlon, J. E. and Johnson, P. T. 1957. On Some Microtine-Infesting Polyplax (Anoplura). Proceedings 
of the Entomological Society of Washington, 59(6):279-283. 
 
Johnson, P. T, 1958. Type Specimens of Lice (Order Anoplura) in the United States National Museum. 
Proceedings of the U. S. National Museum, 108(3393):39-49. 
 
Johnson, P. T. 1958. A New Species of Anoplura from the Philippines. Bulletin of the Brooklyn 
Entomological Society, 53(3):76-80. 
 
Johnson, P. T. 1959. The Rodent-Infesting Anoplura (Sucking Lice) of Thailand with Remarks on Some 
Related Species. Proceedings of the U. S. National Museum, 110(3421):569-597. 
 
Johnson, P. T. 1960. A New Species of Hoplopleura from Australia (Anoplura: Hoplopleuridae). 
Proceedings of the Entomological Society of Washington, 62(2):111-113. 
 
Johnson, P. T. 1962. Notes and Descriptions of African Lice (Anoplura). Proceedings of the 
Entomological Society of Washington, 64(1):51-56. 
 
Johnson, P. T. 1962. Redescriptions of Two Cervid-Infesting Anoplura from Southeast Asia. 
Proceedings of the Entomological Society of Washington, 64(2):107-110. 
 
Johnson, P. T. 1962. The Species of Fahrenholzia Kellog and Ferris from Spiny Pocket Mice (Anoplura: 
Hoplopleuridae). Annals of the Entomological Society of America, 55(4):415-428. 
 
Johnson, P. T. 1962. Three New Anoplura from African Rodents (Anoplura: Hoplopleuridae). 
Proceedings of the Entomological Society of Washington, 64(3):155-165. 
 
Johnson, P. T. 1963. Two Rare Anoplura from Kenya. Proceedings of the Entomological Society of 
Washington, 65(3):226-229. 
 
Johnson, P. T. 1964. The Hoplopleurid Lice of the Indo-Malayan Subregion (Anoplura: Hoplopleuridae). 
Miscellaneous Publications of the Entomological Society of America, 4(3):67-102. 
 
Wenzel, R. L. and Johnson, P. T. 1966. Checklist of the Sucking Lice of Panama (Anoplura). From 
Ectoparasites of Panama, Wenzel, R. L. and Tipton, V. J., (Eds.), pp. 273-279. 
 
Johnson, P. T. 1969. Hamophthirius galeopitheci Mjoberg Rediscovered; with the Description of a New 
Family of Sucking Lice. Proceedings of the Entomological Society of Washington, 71(3):420-428. 
 
Johnson, P. T. 1972. Neohaematopinus appressus, A New Species of Sucking Louse from an Asian Tree 
Squirrel (Anoplura). Pacific Insects, 14(2):389-392. 
 
Johnson, P. T. 1972. Polyplax rhizomydis, A New Species of Sucking Louse (Anoplura) from Asian 
Bamboo Rats. Pacific Insects, 14(2):393-397. 
 
Johnson, P. T. 1972. Hoplopleura intermedia Kellogg and Ferris and Its Allies, with the Description of a 
New Species (Anoplura: Hoplopleuridae). Proceedings of the Entomological Society of Washington, 
74(3):330-337. 
 
Johnson, P. T. 1972. On the Rodent-Infesting Anoplura of Panama. Great Basin Naturalist, 32(3):121-
136. 
 
Johnson, P. T. 1972. Sucking Lice of Venezuelan Rodents, with Remarks on Related Species (Anoplura). 
Brigham Young University Science Bulletin (Biological Series), 17(5):1-62. 
 
Johnson, P. T. 1972. Two New Species of Hoplopleura Enderlein from Laotian Murids (Anoplura). 
Pacific Insects, 14(3):607-611. 
 
 
COLEOPTERA PUBLICATIONS 
 
Alfieri, A. 1976. The Coleoptera of Egypt. Memoires de la Societe Entomologique d’Egypte, Vol. 5, 
1976 361 pp. 
 
Anderson, W. H. 1947. A Terminology For The Anatomical Characters Useful In The Taxonomy of 
Weevil Larvae Proc. Entomol. Soc. Of Wash. 1947, vol. 49(5), pp.123-132 
 
Anderson, D. M. 1956. Notes on Cleonus pPiger (Scop.) in the United States (Coleoptera: 
Curculionoidea). The Coleopterists Bulletin.1956, vol. 10(6), pp.81-86 
 
Anderson, D. M. 1962. The Weevil Genus Smicronyx in America North of Mexico (Coleoptera: 
Curculionoidea). Proc. Of The United States National Museum 1962, vol. 113(3456), pp.185-372 
 
Anderson, D. M. 1963. Notes on the Life History of Anthonomus sycophanta Walsh (Coleoptera: 
Curculionoidea) with Descriptions of the Immature Stages. The Coleopterists Bulletin 1963, vol. 
17(3),pp.73-78 
 
Anderson, D. M.1964. A Review of the Specific Names in North American Miarus (Coleoptera: 
Curculionoidea). The Coleopterists Bulletin vol. 18(1), pp.21-24 
 
Anderson, D. M. 1966. A Simple Photographic Aid to the Illustration of Insects and Other Small 
Organisms. BioScience 1966, vol. 16(10), pp.758-759 
 
Anderson, D. M. 1968. Observations on the Pupae of Anthonomus grandis grandis Boheman and A. 
grandis thurberiae Pierce (Coleoptera: Curculionoidea). Annals of the Entomol. Soc. Of America 1968, 
vol. 61(1), pp.125-129 
 
Anderson, D. M. 1970. Dodder Weevils in Simultaneous Association with Parasitic Plants and Their 
Hosts. Science 1970, vol. 168, pp.132-133 
 
Anderson, D. M. 1970. A Review of the Known Larvae and Pupae Of the Genus Phrydiuchus 
(Coleoptera: Circulionoidea). Annals of the Entomological Society of America 1970, vol. 63(1), pp.296-
300 
 
Anderson, D. M. 1971. Observations On the Larva Of Hypera eximia (LeConte) (Coleoptera: 
Curculionidae). Proceedings of the Entomological Society of Washington 1971, vol. 73(2), pp.117-119 
 
Anderson, D. M. 1971. Oligocaris Lea A Synonym of Smicronyx Schoenherr. Proceedings of the 
Entomological Society of Washington 1971, vol. 73(1), pp.63 
 
Anderson, W. H. and D. M. Anderson. 1971.Type Specimens in the Hans Eggers Collection of Scolytid 
Beetles (Coleoptera). (Smithsonian Contributions to Zoology) 1971, vol. 94 
 
Anderson, D. M. 1973. Keys To Larvae And Pupae of the Gymnetrinae of America North of Mexico 
(Coleoptera: Curculionoidea). Proceedings of the Entomological Society of Washington 1973, vol. 75(2), 
pp.133-140 
 
Anderson, D. M. and W. H. Anderson. 1973. A Key to Separate Larvae Of the Whitefringed Beetles, 
Graphognathus Species, from Larvae of Closely Related Species (Coleoptera: Curculionidae). (U.S. 
Dept. Agr. Coop. Econ. Ins. Rpt.) 1973, vol. 29(49-52), pp.797-800 
 
Anderson, D. M.1974. First Record of Xyleborus semiopacus in the Continental United States 
(Coleoptera:Scolytidae). (U.S. Dept. Agr. Coop. Econ. Ins. Rpt.)1974, vol. 24(45-48), pp.863-864 
 
Anderson, D. M. 1974. Some Species Of Smicronyx (Coleoptera:Circulionoidea) Associated with 
Cuscuta Species (Convolvulaceae) In Pakistan Proceedings of the Entomological Society of Washington 
1974.vol. 76(4), pp.359-374 
 
Perry, R. H., R. W. Surdick, and D. M. Anderson. 1974. Observations on The Biology, Ecology, 
Behavior, And Larvae of Dryobius sexnotatus Linsley (Coleoptera: Cerambycidae). The Coleopterists 
Bulletin 1974, vol. 28(4),pp.169-176 
 
Burke, H. R. and D. M. Anderson. 1976. Systematics of Larvae and Pupae of American Curculionoidea: 
Status Report, Historical Review and Bibliography. The Southwestern Entomologist. 1976, vol.1(2), 
pp.56-73 
 
Clark, W. E., H. R. Burke and D. M. Anderson.1978. The Genus Tychius Germar (Coleoptera: 
Curculionidae): Larvae and Pupae of Some Species, With Evaluation of Their Characters In Phylogenetic 
Analysis. Proceedings of the Entomological Society of Washington 1978, vol. 80(4), pp.626-654 
 
Anderson, D. M. and P. E. Boldt.1979. Identification of Larvae of Two European Species of 
Ceutorhynchinae (Coleoptera: Curculionoidea) Found In Cardus macrocephalus Desfontaines 
(Compositae). Proceedings of the Entomological Society of Washington 1979, vol. 81(3), pp.460-464 
 
Gordon, R. D. and D. M. Anderson. 1979. The Genus Stethorus Weise (Coleoptera: Coccinellidae) in 
Chile. The Coleopterists Bulletin. 1979 vol. 33(1), pp.61-68 
 
Clark, W. E. and D. M. Anderson. 1981. The Genus Chionanthobius Pierce (Coleoptera: Curculionoidea): 
Descriptions of a New Species On Forestiera (Oleaceae) and of the Larva and Pupa of C. schwarzi 
Pierce. Proceedings of the Entomological Society of Washington. 1981,vol. 83(2), pp.185-197 
 
Gordon, R. D. and D. M. Anderson. 1981. The Species of Scarabaeidae (Coleoptera) Associated with 
Sugarcane in South Florida. Florida Entomologist. 1981, vol. 64(1), pp.119-137 
 
McClay, A. S. and D. M. Anderson. 1985. Biology and Immature Stages of Thecesternus hirsutus Pierce 
(Coleoptera: Curculionoidea) in North Eastern Mexico. Proceedings of the Entomological Society of 
Washington 1985, vol. 87(1), pp.207-215 
 
Blackman, M. W. 1931. Black Hill Beetle (Dendroctonus ponderosae). Bull. of N. Y. State College of 
Forestry, Vol. 4, No. 4. 
 
Blake, D. H. 1966. Review of Genus Neobrotica and Some Closely Related Genera (Chrysomelidae). 
Proceedings of the U. S. National Museum, Vol. 118, No. 3529. 
 
Blake, D. H. 1967. Revision of Genus Glyptoscelis (Chrysomelidae). Proceedings of the U. S. National 
Museum, Vol. 123, No. 3604. 
 
Blake, D. H. 1970. Notes on Some Crysomelid Beetles from U. S. A. and Argentina. Proceedings of the 
Entomological Society of Washington, Vol. 172, No. 3. 
 
Blake, D. H. 1976. Brown Semicostate and Costate Species of Colaspis in Mexico and Central America 
(Chrysomelidae). USDA Tech. Bull. No. 1534. 
 
Buchanan, L. L. 1947. A correction and two new races in Graphognathus (white-fringed beetles) 
(Coleoptera: Curculionidae). Journal of the Washington Academy of Sciences, Vol. 37, No. 1: 19-22. 
 
Cartwright and Chalumeau. 1978. Bredin - Archbold - Smithsonian Biological Survey of Dominica 
(Scarabaeoidea). 
 
Gordon and Cartwright. 1988. North American Representatives of Tribe Aegialiini (Scarabaeidae: 
Aphodiinae), SI Contribution to Zoology, No. 461. 
 
Casey, T. L. Misc. Tenebrionidae Papers. 
 
Casey, T. L. 1907. Revision of the American components of the Tenebrionid subfamily Tentyrinae. 
Proceedings of the Washington Academy of Science 9:275-522. 
 
Casey, T. L. 1908. Revision of the Tenebrionid subfamily Coniontinae. Proceedings of the Washington 
Academy of Science 10:51-166. 
 
Casey, T. L. 1916. Memoirs of the Coleoptera. VII. 
 
Erwin, T. L. 1965. A Revision of Brachinus of North America: Part I: The California Species 
(Coleoptera: Carabidae). Coleopterists’ Bulletin, 19(1): 1-19 [17 figs.]. 
 
Erwin, T. L. 1967. Bombardier Beetles (Coleoptera: Carabidae) of North America: Part II: Biology and 
Behavior of Brachinus pallidus Erwin in California. Coleopterists’ Bulletin, 21(2): 41-55 [15 figs.]. 
 
Ball, G. E. and Erwin, T. L. 1969. A Taxonomic Synopsis of the Tribe Loricerini (Coleoptera: 
Carabidae). Canadadian Journal of Zoology, 47(5): 877-907 [62 figs.; 4 tables]. 
 
Erwin, T. L. 1970. A Reclassification of Bombardier Beetles and a Taxonomic Revision of the North and 
Middle American Species (Carabidae: Brachinida). Quaestiones Entomologicae, 6: 4-215 [460 figs.; 16 
tables]. 
 
Erwin, T. L. 1970. A Description of the Larva of Thyce harfordi Casey (Scarabaeidae: Melolonthini). 
Psyche, 77(1): 50-53 [10 figs.]. 
 
Erwin, T. L. 1971. Notes and Corrections to a Reclassification of Bombardier beetles (Carabidae: 
Brachinida). Quaestiones Entomologicae, 7: 281. 
 
Erwin, T. L. and Erwin, J. L. 1971. A Guide to the Ground-Beetle Collection in the Muséum National 
d'Histoire Naturelle, Paris (Coleoptera: Carabidae). Multilithed by SI Duplicating Unit for limited 
distribution, 1-22. 
 
Erwin, T. L. 1972. The Carabid Beetles of New Guinea. Systematic Zoology, 21(3): 343-347. 
 
Erwin, T. L. 1973. Studies of the Subtribe Tachyina (Coleoptera: Carabidae: Bembidiini), Part I: A 
Revision of the Neotropical Genus Xystosomus Schaum. Smithsonian Contributions to Zoology, 140: 1-39 
[72 figs.]. 
 
Erwin, T. L. 1973. A Supplement to the Bombardier Beetles of North and Middle America: New Records 
for Middle America (Coleoptera: Carabidae). Coleopterists’ Bulletin, 27(2): 79-82 [4 figs.]. 
 
Erwin, T. L. 1974. Studies of the Subtribe Tachyina (Coleoptera: Carabidae: Bembidiini), Part II: A 
Revision of the New World-Australian Genus Pericompsus LeConte. Smithsonian Contributions to 
Zoology, 162: 1-96 [161 figs.; 1 table]. 
 
Erwin, T. L. 1974. Studies of the Subtribe Tachyina (Coleoptera: Carabidae: Bembidiini) Supplement A: 
Lectotype Designations for New World Species, Two New Genera, and Notes on Generic Concepts. 
Proceedings of the Entomological Society of Washington, 76(2): 123-155 [2 figs.]. 
 
Erwin, T. L. 1974. The Genus Coptocarpus Chaudoir of the Australian Region with Notes on Related 
African Species (Coleoptera: Carabidae: Oodini). Smithsonian Contributions to Zoology, 184: 1-25 [33 
figs.]. 
 
Erwin, T. L. 1975. Studies of the Subtribe Tachyina (Coleoptera: Carabidae: Bembidiini), Part III: 
Systematics, Phylogeny, and Zoogeography of the Genus Tachyta Kirby. Smithsonian Contributions to 
Zoology, 208: 1-68 [175 figs.]. 
 
Erwin, T. L. 1976. Application of Computerized Information Management Systems (SELGEM) to a 
Natural History Data Bank (Coleoptera: Carabidae). The Coleopterists Bulletin, 30(1):1-31. 
 
Erwin, T. L. 1977. Studies of the Subtribe Tachyina (Coleoptera: Carabidae: Bembidiini) Supplement B: 
Lectotype Designation of Masoreus ridiculus Schaufuss and Transfer of this Species to Polyderis. 
Coleopterists’ Bulletin, 31(1): 90. 
 
Erwin, T. L. 1978. Studies of the Subtribe Tachyina (Coleoptera: Carabidae: Bembidiini) Supplement C: 
Discovery of the Sister Group of Mioptachys Bates, in the Amazon Basin--Inpa psydroides a New Genus 
and Species. Coleopterists’ Bulletin, 32(1): 29-36 [1 fig.]. 
 
Erwin, T. L. 1978. The Larva of Neotropical Enceladus gigas Bonelli (Coleoptera: Carabidae: 
Siagoninae: Enceladini) with notes on the Phylogeny and Classification of some of the More Primitave 
Tribes of Ground Beetles. Coleopterists’ Bulletin, 32(2): 99-106 [4 figs.; 2 tables]. 
 
Erwin, T. L. 1978. Systematic, Natural History, and Zoogeographic Notes on the Genus Agra Fabricius, 
with a Description of a New Species from Panama (Coleoptera: Carabidae: Lebiini). Coleopterists’ 
Bulletin, 32(4): 261-268 [4 figs.]. 
 
Erwin, T. L. and Halpern, A. L. 1978. Max Bänninger: His Collection and Publications (Coleoptera: 
Carabidae). Coleopterists’ Bulletin, 32(4): 357-366 [1 fig.]. 
 
Erwin, T. L. 1979. The American Connection, Past and Present, as a Model Blending Dispersal and 
Vicariance in the Study of Biogeography. pages 355-367 [3 figs.; 2 tables]. in: Carabid Beetles: Their 
Evolution, Natural History, and Classification; Proceedings of the First International Symposium of 
Carabidology. Erwin, T. L., Ball, G. E., and Whitehead, D. R., (Eds.), Dr. W. Junk Publ., The Hague, 
635 pp., . 
 
Erwin, T. L. 1979. A Review of Natural History andEvolution of Ectoparasitoid Relationships in 
Carabid Beetles. pages 479-484 [2 tables]. in: Carabid Beetles: Their Evolution, Natural History, and 
Classification. Proceedings of the First International Symposium of Carabidology. 
 
Erwin, T. L., Ball, G. E., and Whitehead, D. R., (Eds.), Dr. W. Junk Publ., The Hague, 635 pp., . 
 
Erwin, T. L. 1979. Thoughts on the Evolutionary History of Ground Beetles: Hypotheses Generated from 
Comparative Faunal Analyses of Lowland Forest Sites in Temperate and Tropical Regions. pages 539-
592 [19 figs.; 5 tables].in: Carabid Beetles: Their Evolution, Natural History, and Classification. 
Proceedings of the First International Symposium of Carabidology. Erwin, T. L., Ball, G. E., and 
Whitehead, D. R., (Eds.), Dr. W. Junk Publ., The Hague, 635 pp. 
 
Erwin, T. L. 1980. Systematic and Zoogeographic Notes on Apotomini, An Old World Faunal Element 
New to the Western Hemisphere List, With A Description of a New Species From Brazil (Coleoptera: 
Carabidae). Coleopterists’ Bulletin, 34(1): 99-104 [3 figs.]. 
 
Erwin, T. L. and Kavanaugh, D. H. 1980. On the Identity of Bembidion puritanum Hayward (Coleoptera: 
Carabidae: Bembidiini). Coleopterists’ Bulletin, 34(2): 241-242. 
 
Erwin, T. L. and Scott, J. C. 1981. Seasonal and Size Patterns, Trophic Structure, and Richness of 
Coleoptera in the Tropical Arboreal Ecosystem: The Fauna of the Tree Luehea seemannii Triana and 
Planch in the Canal Zone of Panama. Coleopterists’ Bulletin, 34(3): 305-322 
 
Erwin, T. L. and Kavanaugh, D. L. 1981. Systematics and Zoogeography of Bembidion Latreille: I. The 
Carlhi and Erasum Groups of Western North America (Coleoptera: Carabidae: Bembidiini). 
Entomologica Scandinavica Supplement, 15: 33-72 [29 figs.]. 
 
Erwin, T. L. 1981. A Synopsis of the Immature Stages of Pseudomorphini (Coleoptera: Carabidae) with 
notes on Tribal Affinities and Behavior in Relation to Life with Ants. Coleopterists’ Bulletin, 35(1): 53-
68 [29 figs.]. 
 
Erwin, T. L. 1981. Taxon Pulses, Vicariance, and Dispersal: An Evolutionary Syntheses Illustrated by 
Carabid Beetles. pages 159-183. in: Vicariance Biogeography: A Critique. Nelson, G. and Rose, D. E., 
(Eds.), Columbia University Press, New York, . 
 
Erwin, T. L. 1981. Natural History of Plummers Island, Maryland. XXVI. The Ground Beetles of a 
Temperate Forest Site (Coleoptera: Carabidae): An Analysis of Fauna in Relation to Size, Habitat 
Selection, Vagility, Seasonality, and Extinction. Bulletin of the Biological Society of Washington, 5: 105-
224 [34 figs.]. 
 
Erwin, T. L. 1982. Agra, Arboreal Beetles of Neotropical Forests: Erythropus Group Systematics 
(Carabidae). Systematic Entomology, 7: 39-71 [123 figs.]. 
 
Erwin, T. L. 1982. Agra, Arboreal Beetles of Neotropical Forests: Platyscelis-Group Systematics 
(Carabidae). Systematic Entomology, 7: 185-210. 
 
Erwin, T. L. 1982. Small Terrestrial Ground-Beetles of Central America (Carabidae: Bembidiina and 
Anillina). Proceedings of the California Academy of Sciences, 42(19): 455-496 [74 figs; 22 tables]. 
 
Erwin, T. L. 1983. Chaudoir's Studies of the Taxa Agrina and Brachinitae (Coleoptera: Carabidae). 
Coleopterists’ Bulletin, 36(3): 502-512 [4 figs.; 2 tables]. 
 
Ball, G. E. and Erwin, T. L. 1983. The Baron Maximilien de Chaudoir: Inheritance, Associates, Travels, 
Work, and Legacy. Coleopterists’ Bulletin, 36(3): 475-501 [8 figs.; 5 tables]. 
 
Erwin, T. L. 1983. Agra, Aboreal Beetles of Neotropical Forests: Famula Group and Formicaria Groups 
Systematics (Carabidae). Systematic Entomology, 8: 263-292 [75 figs.; 6 tables]. 
 
Erwin, T. L. 1984. Agra, Arboreal Beetles of Neotropical Forests: Palmata-Group Systematics 
(Carabidae). Systematic Entomology, 9: 9-48 [145 figs; 6 tables]. 
 
Erwin, T. L. 1984. Studies of the Tribe Bembidiini (Coleoptera: Carabidae): Lectotype Designations and 
Species Group Assignments for Bembidion Species Described by Thomas L. Casey and others. Pan-
Pacific Entomologist, 60(3): 165-187. 
 
Erwin, T. L. and Sims, L. L. 1984. Carabid Beetles of the West Indies (Insects: Coleoptera): A Synopsis 
of the Genera and Checklists of the Tribes of Caraboidea, and of the West Indian Species. Quaestiones 
Entomologicae, 20(4): 351-466 [2 figs.]. 
 
Erwin, T. L. 1984. Small Terrestrial Ground Beetles of the Amazon Basin (Coleoptera: Bembidiini: 
Tachyina and Anillina). Amazoniana, 8(4): 511-518 [6 figs.]. 
 
Erwin, T. L., 1985. The Taxon Pulse: A General Pattern of Lineage Radiation and Extinction Among 
Carabid Beetles. pages 437-372 [10 figs.]. in: Taxonomy, Phylogeny, and Zoogeography of Beetles and 
Ants. Ball, G. E. (Ed.), Dr. W. Junk Publishers, Dordrecht, . 
 
Kavanaugh, D. K. and Erwin, T. L. 1985. Trechus obtusus Erichson (Coleoptera: Carabidae), a European 
Ground Beetle, on the Pacific Coast of North America: its Distribution, Introduction and Spread. Pan-
Pacific Entomologist, 61(2): 170-179 [3 figs.]. 
 
Erwin, T. L. 1986. [Abstract] Species Diversity in Insects. 33rd Annual Systematics Symposium, 
Missouri Botanical Gardens, St. Louis, MO, 1. 
 
Adis, J. , Paarmann, W. and Erwin, T. L. 1986. On the Natural History and Ecology of Small Terrestrial 
Ground-Beetles (Col.: Bembidiini: Tachyina: Polyderis) from an Amazonian Black-Water Inundation 
Forest. pages 413-427 [6 figs.; 2 tables]. in: Carabid Beetles. den Boer et al., (Eds.), Gustav Fischer, 
Stuttgart/New York, . 
 
Erwin, T. L. 1987. Agra, Arboreal Beetles of Neotropical Forests: Feisthameli Group Systematics 
(Carabidae). Systematic Entomology, 12: 137-161 [55 figs.; 6 tables]. 
 
Erwin, T. L. and Pogue, M. 1988. Agra, Arboreal Beetles of Neotropical Forests: Biogeography and the 
Forest Refugium Hypothesis (Carabidae). pages 161-188 [16 figs.]. in: Proceedings of a Workshop on 
Neotropical Distribution Patterns. Heyer, W. R. and Vanzolini, P. E., (Eds.), Academia Brasileira de 
Ciências, Rio de Janeiro, . 
 
Farrell, B. D. and Erwin, T. L. 1988. Leaf-Beetle Community Structure in an Amazonian Rain Forest 
Canopy. pages 73-90 [6 figs.; 2 tables]. in: The Biology of Chrysomelidae. Jolivet, P., Petitpierre, E. and 
Hsiao, T., (Eds.), Kluwer Academic Publishers, [Place not given], . 
 
Erwin, T. L. 1989. Canopy Arthropod Biodiversity: A Chronology of Sampling Techniques and Results. 
Revista Peruana de Entomologia, 32: 71-77 [5 figs.; 1 table]. 
 
Erwin, T. L. 1991. The Ground-Beetles of Central America (Carabidae) Part II: Notiophilini, Loricerini, 
Carabini. Smithsonian Contributions to Zoology, 501: 1-30 [22 figs.]. 
 
Kavanaugh, D. L. and Erwin, T. L. 1991. The Tribe Cicindini Bänninger (Coleoptera: Carabidae): 
Comparative Morphology, Classification, Natural History, and Evolution. Proceedings of the 
Entomological Society of Washington, 93(2): 356-389 [35 figs.]. 
 
Erwin, T. L. 1991. An Evolutionary Basis for Conservation Strategies. Science, 253: 750-752 [3 figs.]. 
 
Erwin, T. L. 1991. How Many Species Are There?: Revisited. Conservation Biology, 5(3): 330-333 [1 
table]. 
 
Erwin, T. L. 1991. Establishing a Tropical Species Co-occurence Database. Part 1. A Plan for 
Developing Consistent Biotic Inventories in Temperate and Tropical Habitats. Memoiras del Museo de 
Historia Natural, San Marcos, 20: 1-16 [5 figs.]. 
 
Erwin, T. L. and Kable, M. 1991. Establishing a Tropical Species Co-occurence Database. Part 2. An 
Automated System for Mapping Dominant Vegetation. Memorias del Museo de Historia Natural, San 
Marcos, 20: 17-36 [13 figs.]. 
 
Erwin, T. L. and Pearson, D. L. 1991. Establishing a Tropical Species Co-occurrence Database. Part 3. 
An Integrated Approach Toward Understanding Biological Diversity. Memorias del Museo de Historia 
Natural, San Marcos, 20: 37-45 [1 fig.]. 
 
Erwin, T. L. 1991. Natural History of the Carabid Beetles at the BIOLAT Rio Manu, Pakitza, Peru. 
Revista Peruana de Entomologia,33: 1-85 [64 figs.; 3 tables]. 
 
Erwin, T. L. 1994. Arboreal Beetles of Tropical Forests: the Xystosomi Group, Subtribe Xystosomina 
(Coleoptera: Carabidae: Bembidiini) Part I. Character Analysis, Taxonomy, Evolution. Canadian 
Entomologist, 126: 549-666 [155 figs.; 2 tables]. 
 
Erwin, T. L. 1998. Evolution at the equator: aboreal and alticolous beetles and their taxon pulses with 
descriptions of a new Agra subclade and its species (Coleoptera: Carabidae: Lebiini), Mus. reg. Sci. nat. 
Torino, 491-510. 
 
Erwin. T. L. 2000. Arboreal Beetles of Neotropical Forests: Agra Fabricius, the Novaurora Complex 
(Coleoptera: Carabidae: Lebiini: Agrina). Smithsonian Contribution to Zoology, No. 6. 
 
Erwin, T. L. 2000. Arboreal beetles of Neotropical Forests: Taxonomic supplement for the platyscelis 
group with new species and distribution records (Coleoptera:Carabidae, Lebiini, Agrina). The 
Coleopterists Bulletin 54(1):90-119. 
 
Erwin, T. L. 2000. Arboreal beetles of Neotropical Forests: Taxonomic supplement for the Agra virgata 
and ohausi groups with a new species and additional distribution records (Coleoptera:Carabidae). The 
Coleopterists Bulletin 54(2):251-262 
 
Erwin, T. L. and P.J. Johnson. 2000. Naming species, a new paradigm for crisis management in 
taxonomy: Rapid journal validation of scientific names enhanced with more complete descriptions on the 
internet. The Coleopterists Bulletin 54(3):269-278. 
 
Erwin, T. L. 2000. A new genus and species of Lachnophorini and two new species of Lebiini from Costa 
Rica (Coleoptera: Carabidae). The Coleopterists Bulletin 54(3):279-283. 
 
Arndt, E., S. Kimse, and T.L. Erwin. 2001. Arboreal beetles of Neotropical forests: Agra Fabricius, larval 
descriptions with notes on natural history and behaviour (Coleoptera, Carabidae, Lebiini, Agrina). The 
Coleopterists Bulletin 55(3):297-311. 
 
Erwin, T. L. 2002. The Beetle Family Carabidae of Costa Rica: Twenty-nine new species of Costa Rican 
Agra Fabricius 1801 (Coleoptera:Carabidae, Lebiini, Agrina). Zootaxa 119:1-68. 
 
Erwin T. L. and R. F. Medina. 2003. The Neotropical species Askalaphium depressum (Bates): Larval 
description, first diagnosis and illustrations of immature Ctenodactylini, with natural history notes on the 
genus and tribe (Coleoptera: Carabidae). Zootaxa 273:1-15. 
 
Erwin T. L. 2004. The Beetle Family Carabidae of Costa Rica and Panamá: Descriptions of four new 
genera and six new species with notes on their way of life (Insecta: Coleoptera). Zootaxa 537:1-18. 
 
Erwin T. L. 2004. The Beetle Family Carabidae of Costa Rica: The Genera of the Cryptobatida Group of 
Subtribe Agrina, Tribe Lebiini, with new species and notes on their way of life (Insecta: Coleoptera). 
Zootaxa, 662: 1-54. 
 
Erwin T. L. 2004. The Beetle Family Carabidae of Costa Rica: The Genus Epikastea Liebke of the 
Plochonida Group, with new Neotropical species and notes on their way of life (Insecta: Coleoptera, 
Lebiini, Agrina). Zootaxa, 790: 1-20. 
 
Erwin, T. L. and W. Moore. 2007. The Beetle Family Carabidae of Costa Rica: The Genus Moriosomus 
Motschulsky with a new Neotropical species and notes on their way of life (Insecta: Coleoptera, 
Carabidae, Moronini). Zootaxa, 1438: 49-63. 
 
Gerberg, E. J. 1957. Revision of New World Species of Powder-Post Beetles (Lyctidae). USDA Tech. 
Bull. No. 1157. 
 
Kingsolver, J. M. 1974. Biosystematics of Central American Species of Ctenocolum, a New Genus of 
Seed Beetles (Coleoptera: Bruchidae). (Kingsolver and Whitehead, D. R.) 
 
Kingsolver, J. M. 1976. North and Central American Species of Meibomeus (Bruchidae). USDA Tech. 
Bull. No. 1523 
 
Lindroth, C. H. 1979. The Importance of Beringia as Reflected in the Present Fauna. In: Carabid 
Beetles: Their Evolution, Natural History, and Classification. Proceedings of the First International 
Symposium of Carabidology. Erwin, T. L., Ball, G. E., and Whitehead, D. R., (Eds.), Dr. W. Junk Publ., 
The Hague, 635 pp. 
 
Lindroth, C. H. 1979. South American Carabid Fauna: Endemic Tribes and Tribes with African 
Relationships. In: Carabid Beetles: Their Evolution, Natural History, and Classification. Proceedings of 
the First International Symposium of Carabidology. Erwin, T. L., Ball, G. E., and Whitehead, D. R., 
(Eds.), Dr. W. Junk Publ., The Hague, 635 pp., 
 
Lindroth, C. H. 1979. Europe and the Circumpolar Area: Events and Personalities. In: Carabid Beetles: 
Their Evolution, Natural History, and Classification. Proceedings of the First International Symposium of 
Carabidology. Erwin, T. L., Ball, G. E., and Whitehead, D. R., (Eds.), Dr. W. Junk Publ., The Hague, 
635 pp. 
 
Lindroth, C. H. 1979. Theory of Glacial Refugia. In: Carabid Beetles: Their Evolution, Natural History, 
and Classification. Proceedings of the First International Symposium of Carabidology. Erwin, T. L., Ball, 
G. E., and Whitehead, D. R., (Eds.), Dr. W. Junk Publ., The Hague, 635 pp. 
 
Lindroth, C. H. 1992. Ground Beetles of Fennoscandia, Part I. 
 
Lindroth, C. H. 1992. Ground Beetles of Fennoscandia, Part III. 
 
Menke, A. S., and F. D. Parker. 1988. Adult Feeding and Distribution of Dynastes Granti Horn 
(Coleoptera:Scarabaeidae). The Coleopterists Bulletin 42(2), 1988, pp. 161-164 
 
Pfaffensberger, G. S. and C. D. Johnson. 1976. Biosystematics of First-Stage Larvae of Some North 
American Bruchidae. USDA Tech. Bull. No. 1525. 
 
Spangler, P. J. 1950. Occurrence of Scaphiophus holbrooki holberooki (Harlan) in Athens County, Ohio. 
Ohio Journal of Science 50(6):277. 
 
Spangler, P. J. 1966. Results of Catherwood Foundation Peruvian-Amazon Expedition, Insects, Part XIII, 
The Aquatic Coleoptera Dytiscidae; Noteridae; Gyrinidae; Hydrophilidae; Dascillidae; Helodidae; 
Psehpenidae; Elmidae). Monograph 14, Academy of Natural Sciences Philadelphia, pages 377-443, 129 
figures. 
 
Spangler, P. J. 1968. A New Psephenus and Its Larva from Mexico (Coleoptera:Psephenidae) 
Etomological News 79(4):91-97, 11 figures. 
 
Fales, J. H., P. J. Spangler, O. F. Bodenstein, G. D. Mills, and C. C. Durbin, Jr. 1968. Laboratory and 
Field Evaluations of AbateR, Againsta Backswimmer,Notonecta undulata Say Hemiptera: Notonectidae). 
Mosquito News 28(1):77-81, 5 tables. 
 
Spangler, P. J. 1972. A New Genus and New Species of Water Beetle from Bolivia with a Key to the 
Genera of the Western Hemisphere Colymbetini (Coleoptera: Dytiscidae). Proceedings of the Biological 
Society of Washington 84(49):427-434, 7 figures. 
 
Spangler, P. J. 1972. A New Genus and Two New Species of Madicolous Water Beetles from Venezuela 
(Coleoptera: Hydrophilidae). Proceedings of the Biological Society of Washington 85(10):139-146, 13 
figures. 
 
Spangler, P. J. and J. L. Cross. 1972. A Description of the Egg Case and Larva of the Water Scavenger 
Beetle, Helobata striata (Coleoptera: Hydrophilidae). Proceedings of the Biological Society of 
Washington 85(35):413-418, figures. 
 
Spangler, P. J. and J. M. Gillespie. 1973. The Larva and Pupa of the Predaceous Water Beetle, Hygrotus 
sayi (Coleoptera: Dytiscidae). Proceedings of the Biological Society of Washington 86(12):143-152, 11 
figures. 
 
Spangler, P. J. and R. D. Gordon. 1973. Descriptions of the Larvae of Some Predacious Water Beetles 
(Coleoptera: Dytiscidae). Proceedings of the Biological Society of Washington 86(22):261-278, 16 
figures. 
 
Spangler, P. J. and G. W. Folkerts. 1973. The Larva of Pachydrus princeps (Coleoptera:Dytiscidae). 
Proceedings of the Biological Society of Washington 86(29):351-356, 3 figures. 
 
Spangler, P. J. 1973. Aquatic Coleoptera Collected by the Biospeleological Expeditions to Cuba by the 
Academies of Science of Cuba and Romania. Pages353--358, 4 figures. In: Orghidian et al.; Resultats 
des Expeditions Biospeologiques Cuban-Roumaines a Cuba 1. Bucarest, Romania: Editura Academiei 
Republicii Socialiste Romania. 
 
Spangler,P. J. 1973. The Bionomics, Immature Stages, and Distribution of the Rare Predacious Water 
Beetle, Hoperius planatus (Coleoptera: Dytiscidae). Proceedings of the Biological Society of 
Washington 86(36):423-434, 12 figures. 
 
Spangler, P. J. 1973. The Nomenclature, Bionomics, and Distribution of Notaticus fasciatus 
(Coleoptera: Dytiscidae: Aubehydrinae). Proceedings of the Biological Society of Washington 86(42): 
495-500, 8 figures. 
 
Spangler, P. J. and G. W. Folkerts. 1973. Reassignment of Colpius inflatus and a Description of its 
Larva (Coleoptera: Noteridae). Proceedings of the Biological Society of Washington 86(43): 501-510, 11 
figures. 
 
Spangler, P. J. 1974. A Description of the Larva of Hydrobiomorpha casta (Coleoptera: Hydrophilidae). 
Journal of the Washington Academy of Science 63(4):160--164, 7 figures. [1973]. 
 
Spangler, P. J. 1974. A Description of the Larva of Celina angustata Aubé (Coleoptera: Dytiscidae). 
Journal of the Washington Academy of Science 63(4):165-168, 8 figures. [1973]. 
 
Spangler, P. J. 1974. The Rediscovery of Cylorygmus lineatopunctatus (Coleoptera: Hydrophilidae: 
Sphaeridiinae: Rygmodini). Journal of the Kansas Entomological Society 47(2):244-248, 7 figures. 
 
Reichardt, H., S. Vanin, and P. J. Spangler. 1975. New and Little Known Neotropical Coleoptera IV. 
Notes on Spercheidae, especially Spercheus fimbricollis Bruch. Papeis Avulsos de Zoologia, Sao Paulo 
29(11):71-78, 12 figures. 
 
Spangler, P.J. and P. D. Perkins. 1977. Three New Species of the Neotropical Water Beetle Genus 
Elmoparnus (Coleoptera: Dryopidae). Proceedings of the Biological Society of Washington 89(63):743-
760, 20 figures. 
 
Longley, G. and P. J. Spangler. 1977. The Larva of a New Subterranean Water Beetle, Haideoporus 
texanus (Coleoptera: Dytiscidae: Hydroporinae). Proceedings of the Biological Society of Washington 
90(3):532-535, 4 figures. 
 
Spangler, P. J. 1977. Three New Ecuadorian Species of the Aquatic Beetle Genus Chaetarthria 
(Coleoptera: Hydrophilidae).Proceedings of the Biological Society of Washington 90(3):566-578, 15 
figures. 
 
Spangler, P.J., and S. Santiago. 1977. A New Water Scavenger Beetle from Mexico (Coleoptera: 
Hydrophilidae). Proceedings of the Biological Society of Washington 92(2):333-338, 11figures. 
 
Spangler, P. J. 1979. A New Genus of Water Beetle from Austral South America (Coleoptera: 
Hydrophilidae). Proceedings of the Bio Soc of Washington 92(4):697-718 
 
Spangler, P. J. 1979. Description of the Larva and Pupa of Cylorygmus ineatopunctatus (Coleoptera: 
Hydrophilidae: Rygmodini). Proceedings of the Biological Society of Washington 92(4):743-752, 14 
figures. 
 
Spangler, P. J. 1979. A New Genus of Madicolous Beetle from Ecuador (Coleoptera: Hydrophilidae: 
Hydrobiinae). Proceedings of the Biological Society of Washington 92(4):753-761, 8 figures. 
 
Spangler, P. J. 1979. Hydramara argentina, a Description of its Larva and a Report on its Distribution 
(Coleoptera: Hydrophilidae). Proceedings of the Biological Society of Washington 81(4):536-543, 11 
figures. 
 
Spangler, P. J. 1979. A New Genus of Water Scavenger Beetle from Chile (Coleoptera: Hydrophilidae: 
Sphaeridiini). Coleopterists Bulletin 33(3):303-308. 
 
Spangler, P. J. 1980. A New Species of the Riffle Beetle Genus Portelmis from Ecuador (Coleoptera: 
Elmidae). Proceedings of the Entomological Society of Washington 82(1):63-68. 
 
Spangler, P. J. 1980. Onopelmus, a New Genus of Dryopid Beetle from Peru (Coleoptera: Dryopidae). 
Proceedings of the Entomological Society of Washington 82(2):161-165. 
 
Spangler, P. J. 1980. Two New Species of the Water Beetle Genus Hydraena from Cuba. Proceedings of 
the Entomological Society of Washington 82(2):329-333, 4 figures. 
 
Spangler, P. J. 1980. Chelonariid Larva, Aquatic or Not? Coleopterists Bulletin 34(1):105-114. 
 
Spangler, P. J. 1980. A New Species of Ytu from Brazil (Coleoptera: Torridincolidae). Coleopterists 
Bulletin 34(2):145-158, 20 figures. 
 
Spangler, P. J. 1980. Aquatic Coleoptera. In: S.W. Roback (ed.). The Results of the Catherwood 
Foundation Bolivian-Peruvian Altiplano Expedition. Part I. Aquatic Insects Except Diptera. 
Proceedings of the Acad of Natural Sciences of Philadelphia 132:199-212. 
 
Spangler, P. J. 1980. The Aquatic Beetles Collected on the Starmühlner Mission to Reunion and Comoro 
Islands (Coleoptera: Hydrophilidae, Hydraenidae, Gyrinidae). Ann. Naturhist. Mus. Wien 83:701-709, 4 
figures. 
 
Spangler, P. J. 1981. New and Interesting Water Beetles from Mt. Roraima and Ptari-tepui, Venezuela 
(Coleoptera: Dytiscidae and Hydrophilidae). Aquatic Insects 3(1):1-11, 17 figures. 
 
Spangler, P. J. 1981. A New Water Beetle, Troglochares ashmolei, n. gen., n. p., from Ecuador; the First 
Known Eyeless Cavernicolous Hydrophilid Beetle (Coleoptera: Hydrophilidae). Proceedings of the 
Entomological Society of Washington, 83(2):316-323, 9 figures. 
 
Perkins, P. D., and P. J. Spangler. 1981. A Description of the Larva of Helocombus bifidus (Coleoptera: 
Hydrophilidae). Pan-Pacific Entomologist 57(1):52-56. 
 
Spangler, P. J. 1981. Two New Genera, Two new Species of Bidessine Water Beetles from South 
America (Coleoptera: Dytiscidae). Pan-Pacific Entomologist 57(1):65-75, 12 figures. 
 
Spangler, P. J. and P. D. Perkins. 1981. A New Genus and Species of Water Scavenger Beetle from 
Africa (Coleoptera: Hydrophilidae: Sphaeridiinae). Pan-Pacific Entomologist 57(1):220-226, 4 figures. 
 
Spangler, P. J. 1981. A Durable and Lightweight Insect Net and a Manual Aspirator for Collecting 
Aquatic Organisms. Pan-Pacific Entomologist 57(1):245-250, 4 igures. 
 
Spangler, P. J. 1981. Pagelmis amazonica, a New Genus and Species of Water Beetle from Ecuador 
(Coleoptera: Elmidae). Pan-Pacific Entomologist 57(1):286-294, 17 figures. 
 
Spangler, P. J. 1981. Sphaeriidae in Mollusca and Insecta: Comments on Proposal to Remove the 
Homonymy. Z.N.S)1892 (see Bull. Zool. Nom. vol. 32, pp. 60--62, 201-204). Bulletin of Zoolological 
Nomenclature, 38(3):157--161. 
 
Spangler, P. J. 1981. Coleoptera. In: S.H. Hurlbert, G. Rodriquez, and N. D. Dos Santos (eds.). Aquatic 
Biota of Tropical South America, Part l. Arthropoda. June, 1981:129-220. San Diego State University, 
San Diego, California. 
 
Spangler, P. J., and H. P. Brown, H. P. 1981. Discovery of Hydora a Hitherto Australian-New Zealand 
Genus of Riffle Beetle in South America (Coleoptera: lmidae). Proceedings of the Entomological Society 
of Washington, 83(4):596-606, 5figures. 
 
Cartwright, O. L. and P. J. Spangler. 1981. A New Ataenius from the Socorro Islands, Mexico 
(Coleoptera; Scarabaeidae; Aphodiinae). Proceedings of the Entomological Society of Washington 
83(4):785-789, 5 figures. 
 
Spangler, P. J. 1981. Two New Genera of Phreatic Elmid Beetles from Haiti; One Eyeless and One with 
Reduced Eyes (Coleoptera: Elmidae). Bijdragen tot de Dierkunde (Contributions to Zoology). 
(Amsterdam) 51(2):375-387, 44 figures. 
 
Spangler, P. J. 1981. Supplement to the Aquatic and Semiaquatic Coleoptera of Cuba Collected by the 
Biospeleological Expeditions to Cuba by the cademies of Science of Cuba and Romania; pages 145-171, 
8 figures. In: Résultats des Expéditions Biospéologiques Cubano-Roumaines à Cuba 3. Editura 
Academie Republicii Socialiste România / Bucuresti. 
 
Spangler, P. J., and A. Huacuja. 1982. Deltostethus scitulus, a New Hydrophilid Beetle from Mexico 
(Coleoptera: Hydrophilidae: Sphaeridiinae). Entomological News 93(1):1-7, 6 figures. 
 
Spangler, P. J., and A. Vega. 1982. The First Record of Hydrovatus horni Crotch from the Antilles with 
Notes on its Known Distribution and Status (Coleoptera: Dytiscidae). Entomological News 93(2):37-41, 
5 figures. 72. 
 
Spangler, P.J. 1982. Coleoptera; pages 328-397.In: S.H.Hurlbert and A.Villalobos-Figueroa (eds). 
Aquatic Biota of Mexico, Central America, and The West Indies. San Diego State University, San Diego, 
California. 
 
Spangler, P. J., and S. Santiago. 1982. A New Species of Aquatic Beetle, Disersus uncus, from Costa 
Rica (Coleoptera: Elmidae: Larinae). Special Issue to the Memory of Retirement of Emeritus Professor 
Michio Chujo. December, l982:l7-20, 4 figures. 
 
Siebert, H., P. J. Spangler, and F. Stehr. l982. Obituary: William Carl Stehr. Bulletin Of The 
Entomological Society Of America 28(4):380, l figure. 75. 
 
Spangler, P.J., and Steiner, W.E. 1983. New Species of Water Beetles of the Genera Elmoparnus and 
Pheneps from Suriname (Coleoptera: Dryopidae;Psephenidae). Proceedings of the Entomological Society 
of Washington 85(4):826-839, 25 figures. 
 
Spangler, P. J. 1983. Immature Stages and Biology of etraglossa palpalis Champion (Coleoptera: 
Ptilodactylidae). Entomological News 94(5):161-175, 23 figures. 
 
Spangler, P. J. 1984. A Survey of Aquatic Insects of Ecuador. National Geographic Society Research 
Reports (1975). 16:611--14. 
 
Wolfe, G. W., and P. J. Spangler. 1985. A Synopsis of the Laccornis ifformis Species Group with a 
Revised Key to North American Species of Laccornis Des Gozis (Coleoptera: Dytiscidae). Proceedings 
of the Biological Society of Washington 98(1):61-71, 17 figures. 
 
Spangler, P. J. 1985. A New Species of the Aquatic Beetle Genus Dryopomorphus from Borneo 
(Coleoptera: Elmidae: Larinae). Proceedings of the Biological Society of Washington 98(2):416-421, 9 
figures. 
 
Perkins, P.D., and P. J. Spangler. 1985. Quadryops, New Genus, and Three New Species of Arboreal 
Dryopidae (Insecta: Coleoptera) from Panama and Ecuador. Proceedings of the Biological Society of 
Washington 98(2):494-510, 47 figures. 
 
Spangler, P. J. 1985. A New Genus and Species of Riffle Beetle, Neblinagena prima, from the 
Venezuelan tepui, Cerro de a Neblina (Coleoptera, Elmidae, Larinae). Proceedings of the Entomological 
Society of Washington 87(3):538--544, 15 figures. 
 
Spangler, P. J. 1985. Obituary: Oscar Ling Cartwright. Proceedings of the Entomological Society of 
Washington 87(3):690-698, 1 figure. 
 
Spangler, P. J. 1985. Five New Species of the Predacious Water Beetle Genus Hydrodessus from Guyana 
and a Key to the Species (Coleoptera: Dytiscidae). Proceedings of the Academy of Natural Sciences of 
Philadelphia 137:80-89, 26 figures. 
 
Spangler, P. J., R. C. Froeschner, and J. T. Polhemus. 1985. Comments on a Water-strider, 
Rheumatobates meinerti, from the Antilles and a Checklist of the Species of the Genus (Hemiptera: 
Gerridae). Entomological News 96(5):196-200. 
 
Spangler, P. J. 1986. The Status of the Riffle Beetle Genus Lara LeConte and Homonymy of the 
Subfamily Group Name Larinae (Coleoptera: Elmidae). Entomological News 97(2):77-79. 
 
Spangler, P. J. 1986. Insecta: Coleoptera. pages 622--631, 16 figures. In: Botosaneanu, L. (ed). 
Stygofauna Mundi. A Faunistic, Distributional, and Ecological Synthesis of the World Fauna Inhabiting 
Subterranean Waters (Including the Marine Interstitial). Leiden:E.J. Brill/Dr. W. B ackhuys. 
 
Spangler, P. J. 1986. Two New Species of Water-striders in the Genus Oiovelia from the tepui Cerro de 
la Neblina, Venezuela (Hemiptera: Veliidae). Proceedings Entomological Society of Washington 
88(3):438-450, 26 figures. 
 
Spangler, P. J. 1986. A New Genus and Species of Water Scavenger Beetle, Guyanobius adocetus, from 
Guyana and Its Larva (Coleoptera: Hydrophilidae: Hydrobiinae). Proceedings Entomological Society of 
Washington 88(3):585-594, 24 figures. 
 
Spangler, P. J. 1986. Three New Species of Water Scavenger Beetles of the Genus Chaetarthria from 
South America (Coleoptera: Hydrophilidae). Proceedings of the Biological Society of Washington 
99(3):509-516, 8 figures. 
 
Spangler, P. J. and S. Santiago. 1986. Una nueva especie de Coleoptero acuatico, del genero Macrelmis 
Motschulsky de Mexico y Centroamerica (Coleoptera: Elmidae). Annals Instituto de Biologia 
Universidad Nacional Autónoma de Mexico 56(ser. zool.)(1):155-158, 3 figures. [1985]. 
 
Spangler, P. J. and P. D. Perkins, P. D. 1986. A New Species of Aquatic Beetle, Elmoparnus jordanorum, 
from Guyana (Coleoptera: Dryopidae). Coleopterists Bulletin 40(4):359-365, 8 figures. 
 
Spangler, P. J. and R. C. Froeschner. 1987. Distributional Data, Illustrations, and Habitat of the South 
American water-strider Microvelia ayacuchana (Hemiptera: Veliidae). Proceedings of the Entomological 
Society of Washington 89(1):167-174, 17 figures. 
 
Spangler, P. J. and S. Santiago. 1987. A Revision of the Neotropical Aquatic Beetle Genera Disersus, 
Pseudodisersus, and Potamophilops (Coleoptera: Elmidae). Smithsonian Contributions to Zoology 46:i-
iii, 1-40, 140 figures. 
 
Spangler, P. J. 1987. Laridae Vigors, 1825 (Aves) and Larini LeConte, 1861 (Insecta, Coleoptera): 
Proposal to Remove the Homonymy. Bulletin of Zoological Nomenclature 44(1):25-26. 
 
Spangler, P. J. 1987. A New Species of Water Penny Beetle, Pheneps cursitatus, from Cerro de la 
Neblina, Venezuela (Coleoptera: Dryopoidea: Psephenidae). Proceedings of the Entomological Society of 
Washington 89(2):219-225, 15 figures. 
 
Spangler, P. J. 1987. Holcodyrops mouli, an Anomalous New Genus and Species of Beetle from Ecuador 
(Coleoptera: Dryopidae). Proceedings of the Entomological Society of Washington 89(3):616-621, 15 
figures. 
 
Spangler, P. J. 1988. Stygobiontic Beetles Collected from Caves in Indonesia (1986) and Thailand 
(1987); pages 21-22. In: Expéditions de l'APS en Asie du Sud-Est, travaus scienti fiques 1. 
 
Spangler, P. J. 1989. A New Species of Neotropical Water Bug, Paravelia biae from Brazil (Heteroptera: 
Veliidae). Proceedings of the Entomological Society of Washington 91(3):360-366, 16 figures. 
 
Polhemus, J. T. and P. J. Spangler. 1989. A New Species of Rheumatobates Bergroth from Ecuador and 
Distribution of the Genus (Heteroptera: Gerridae). Proceedings of the Entomological Society of 
Washington 91(3):421-428, 4 figures. 
 
Spangler, P. J. and P. D. Perkins. 1989. A Revision of the Neotropical Riffle Beetle Genus Stenhelmoides 
(Coleoptera: Elmidae). Smithsonian Contributions to Zoology 479:iv + 63 pages, 262 figures. 
 
Spangler, P. J. 1990. A New Species of Halophilous Water-strider, Mesovelia polhemusi, from Belize 
and a Key and Checklist of New World species of the Genus (Heteroptera: Mesoveliidae). Proceedings 
of the Biological Society of Washington 103(1):86-94, 23 figures. 
 
Spangler, P. J. 1990. A New Species and New Record of the Water-penny Genus Psephenops 
(Coleoptera: Psephenidae) from Costa Rica. Entomological News 101(3):137--140, 4 figures. 
 
Spangler, P. J. 1990. A New Species of Water Scavenger Beetle, Guyanobius simmonsorum from Brazil 
(Coleoptera: Hydrophilidae). Proceedings of the Entomological Society of Washington 92(3):411-415, 13 
Figures. 
 
Spangler, P. J. 1990. A Revision of the Neotropical Aquatic Beetle Genus Stegoelmis (Coleoptera: 
Elmidae). Smithsonian Contributions to Zoology 502:iii + 52 pages, 189 figures. 
 
Spangler, P. J. 1990. Associate Editor, Insects. Index to Proceedings of the Biological Society of 
Washington Volumes 1-100 Inclusive 1880-1987; 193 pages. Compiled by Phyllis Spangler. Editors: 
Phyllis Spangler and C. Brian Robbins. Lawrence, Kansas: Allen Press. 
 
Spangler, P. J. 1991. Haliplidae, pages 311--312, 2 figures. In F.W. Stehr, editor, Immature Insects. 
2:975 pages. Dubuque, Iowa; Kendall/Hunt. 
 
Spangler, P. J. 1991. Noteridae, pages 314-315, 6 figures. In F.W. Stehr, editor, Immature Insects. 2:975 
pages. Dubuque, Iowa; Kendall/Hunt. 
 
Spangler, P. J. 1991. Dytiscidae, pages 315-319 19 figures. In F.W. Stehr, editor, Immature Insects. 
2:975 pages. Dubuque, Iowa;Kendall/Hunt. 
 
Spangler, P. J. 1991. Gyrinidae, pages 319-320, 6 figures. In F.W. Stehr, editor, Immature Insects. 
2:975 pages. Dubuque, Iowa; Kendall/Hunt. 
 
Spangler, P. J. 1991. Hydraenidae, pages 320-322, 6 figures. In F.W. Stehr, editor, Immature Insects. 
2:975 pages. Dubuque, Iowa; Kendall/Hunt. 
 
Spangler, P. J. 1991. Hydrophilidae, pages 355-358, 12 figures. In F.W. Stehr, editor, Immature Insects. 
2:975 pages. Dubuque, Iowa; Kendall/Hunt. 
 
Spangler, P. J. 1991. Georyssidae, pages 358-359, 5 figures. In F.W. Stehr, editor, Immature Insects. 
2:975 pages. Dubuque, Iowa; Kendall/Hunt. 
 
Spangler, P. J. 1991. Chelonariidae, pages 394-395,8 figures. In F. W. Stehr, editor, Immature Insects. 
2:975 pages. Dubuque, Iowa; Kendall/Hunt. 
 
Spangler, P. J., and S. Santiago, S. 1991. A New Species and New Records from Colombia of the Water 
Beetle Genus Onychelmis Hinton (Coleoptera: Elmidae: Elminae). Proceedings of the Entomological 
Society of Washington 93(2):495-498, 4 figures. 
 
Spangler, P. J. and R. A. Faitoute. 1991. A New Genus and Species of Neotropical Water Beetle, 
Jolyelmis auyana, from a Venezuelan Tepui (Coleoptera: Elmidae). Proceedings of the Biological 
Society of Washington 104(2):322-327, 25 figures. 
 
Spangler, P. J. 1991. Guaranius carlosi, a New Genus and Species of Dryopid Beetle from Paraguay 
(Coleoptera: Dryopidae). Coleopterists Bulletin 45(2):147-151, 4 figures. 
 
Barr, C. B. and P. J. Spangler, P. J. 1992. A New Genus and Species of Stygobiontic Dryopid Beetle, 
Stygoparnus comalensis (Coleoptera: Dryopidae), from Comal Springs, Texas. Proceedings of the 
Biological Society of Washington 105(1):40-54, 46 figures. 
 
Spangler, P. J. and S. Santiago-Fragoso. 1992. The Aquatic Beetle Subfamily Larainae (Coleoptera: 
Elmidae) in Mexico, Central America, and the West Indies. Smithsonian Contributions to Zoology 528:vi 
+ 74, 267 figures. 
 
Strand, R. M. and P. J. Spangler. 1994. The Natural History, Distribution, and Larval Description of 
Brychius hungerfordi Spangler (Coleoptera: Haliplidae). Proceedings of the Entomological Society of 
Washington 96(2):208-213, 15 figures. 
 
Barr, C. B. and P. J. Spangler. 1994. Two New Synonymies: Alabameubria, Brown, a Junior Synonym of 
Dicranopselaphus and Alabameubria starki, a Synononym of Dicranopselaphus variegatus (Coleoptera: 
Psephenidae). Entomological News 105(5):299-302, 2 figures. 
 
Polhemus, J. T. and P. J. Spangler, P. J. 1995. A Review of the Genus Stridulivelia Hungerford and Two 
New Species (Heteroptera, Veliidae) from South America. Proceedings of the Entomological Society of 
Washington 97(1):128-152, 20 figures. 
 
Spangler, P. J. 1995. A Review and Two New Species of the Genus Pseudeucinetus Heller from 
Southeast Asia and a World Checklist of the Thaumastodinae (Coleoptera: Limnichidae); pages 395-405, 
35 figures. In, Watanabe,J., Satô, M., and Owada, M. (eds), Beetles and Nature. Special Bulletin of the 
Japanese Society of Coleopterology Number 4, IX + 510. Tokyo. 
 
Spangler, P. J. and C. B. Barr. 1995. A New Genus and Species of Stygobiontic Dytiscid Beetle, 
Comaldessus stygius (Coleoptera: Dytiscidae: Bidessini) from Comal Springs, Texas. Insecta Mundi 9(3 
and 4):301-308, 26 figures. 
 
Spangler, P. J. 1996. A New Genus and Species of Aquatic Beetle, Caenelmis octomeria, from Kenya, 
Africa (Coleoptera: Elmidae: Elminae). Insecta Mundi 10(1-4):19-23, 20 figures. 
 
Spangler, P. J. 1996. Four New Stygobiontic Beetles (Coleoptera: Dytiscidae; Noteridae; Elmidae). 
Insecta Mundi 10(1–4):241-259, 79 figures. 
 
Spangler, P. J. 1997. Two New Species of the Aquatic Beetle Genus Macrelmis Motschulsky from 
Venezuela (Coleoptera: Elmidae: Elminae). Insecta Mundi 11(1):1-8, 25 figures. 
 
Alarie, Y., L-J. Wang, A. N. Nilsson, and P. J. Spangler. 1997. Larval Morphology of Four Genera of 
the Tribe Hyphydrini Sharp (Coleoptera: Dytiscidae) with an Analysis of Their Phylogenetic 
Relationships. Annals of the Entomological Society of America 90(6):709-735, 54 illustrations, 5 tables. 
 
Boobar, L. R., P. J. Spangler, K. E. Gibbs, J. R. Longcore, and K. M. Hopkins. 1998. Predaceous Diving 
Beetles in Maine: Faunal List and Keys to Subfamilies. Northeastern Naturalist 5(1):1-20. 6 plates. 
 
Spangler, P. J. and V. Decu. 1998. The Stygobiontic Coleoptera. In Juberthie, C., and Decu, V. 
Encyclopaedia Biospeologica, Tome 2:1031-1046, 30 figures. 
 
Alarie, Y., P. J. Spangler, P. D. Perkins. 1998. Study of the Larvae of Hydrotrupes palpalis Sharp 
(Coleoptera: adephaga, Dytiscidae) with Implications for the Phylogeny of the Colymbetinae. The 
Coleopterists Bulletin 52(4):313-332. 22 figures. 
 
Spangler, P. J. 1999. A New Tribe, Genus, and Species of Limnichid Beetle, Wooldridgeus perforatus, 
from Malaysia (Coleoptera: Limnichidae: Wooldridgeini). Insecta Mundi 12(3 and 4):181-187. 32 
figures. [1998] 
 
Santiago-Fragoso, S. and P. J. Spangler. 2000. Elmidae (Coleoptera), pages 421-438, 25 figures. In 
Llorente Bousquets, J., González Soriano, E., and Papavero, N. (eds). Biodiversidad, Taxonomía y 
Biogeografía de Artrópodos de México: Hacia una Síntesis de su Conocimiento 2. 676 pages. 
Universidad Nacional Autónoma de México. Ciudad Universitaria / México 
 
Vogt, G. B. 1949. Notes on Cerambycidae from the Lower Rio Grande Valley, Texas. Pan-Pacific Ent. 
25(3):137-144. 
 
Warner, R. miscellaneous publications between 1939-1979 are available (request any desired 
publications). 
 
White, R. E. 1968. Review of the Genus Cryptocephalus in America North of Mexico Coleoptera: 
Chrysomelidae, R. E. White, United States National Museum Bulletin 290. 
 
White, R. E. 1973. Revision of the Genus Petalium Le Conte in the U.S.A., Geater Antilles, and the 
Bahamas (Anobiidae). USDA Tech. Bull. No. 1467. 
 
White, R. E. 1974. Dorcatominae and Tricoryninae of Chile, Coleoptera: Anobiidae. Transactions of the 
American Entomological Society, Vol. 100:191-253. 
 
White, R. E. 1980. Taxonomic Study of the New World Genus Stichtoptychus Fall Coleoptera: 
Anobiidae. 
 
White, R. E. 1980. Taxonomic Study of New World Genus Stichtoptychus Fall (Anobiidae). USDA Tech. 
Bull. No. 1602. 
 
White, R. E. 1981. Homonymy in World Species-Group Names of Criocerinae (Chrysomelidae). USDA 
Tech. Bull. No. 1629. 
 
White, R. E. 1993. Revision of Subfamily Criocerinae (Chrysomelidae) of North America North of 
Mexico. USDA Tech. Bull. No. 1805. 
 
Whitehead, D. R. 1972. Classification, Phylogeny, and Zoogeography of Schizogenius Putzeys 
(Coleoptera: Carabidae). Quaestiones Entomologicae, 8:131-348. 
 
Whitehead, D. R. 1973. Annoted Key To Platynus, Including Mexisphodrus and Most “Colpodes”, so far 
Described from North America Including Mexico (Coleoptera: Carabidae, Quaestiones Entomologicae, 
9:173–217. 
 
Whitehead, D. R. 1974. An Annotated Key to Described Species of the Neotropical Genus Glyptolenus 
(Carabidae: Pterostichini). The Coleopterists Bulletin, 28(3). 
 
Whitehead, D. R. 1975. Classification of the Middle American Genus Cyrtolaus Bates (Coleoptera: 
Carabidae: Pterostichini), Whitehead and Ball, G. E. Quaestiones Entomologicae, 11(4). 
 
Whitehead, D. R. 1975. Megasennius, A New Genus for Acanthoscelides muricatus (Sharp) Coleoptera: 
Bruchidae), A Seed Predator of Cassia grandis L. (Caesalpiniaceae) in Central America. Proceedings of 
the Entomological Society of Washington, Vol. 77, No. 4. 
 
Whitehead, D. R. 1977. Classification of Listropus Putzeys, a Subgenus of Schizogenius Putzeys 
(Coleoptera: Carabidae). (Whitehead and Reichardt). The Coleopterists Bulletin, 31(3). 
 
Whitehead, D. R. 1990. What is an Immigrants Arthropod?, Whitehead and Wheeler, A. G., Jr. Annals of 
the Entomological Society of America, Vol. 83, No. 1 (9-14). 
 
 
USDA CATALOG OF THE COLEOPTERA OF AMERICA NORTHOF MEXICO 
 
Fascicles currently available: 
 
Allocorynidae: C. W. O’Brien, 1991 
Amphizoidae: D. H. Kavanaugh, 1984 
Anobiidae: R. E. White, 1982 
Byturidae: C. A. Springer, 1991 
Cerylonidae: J. F. Lawrence, 1982 
Dryopidae: H. P. Brown, 1983 
Elmidae: H. P. Brown, 1983 
Ciidae: J. F. Lawrence, 1982 
Cupedidae: M. D. Atkins, 1979 
Curculionidae: Acicnemidinae, Cossoninae, Dryophthorinae, Molytinae, Petalochilinae, Rhytirrhininae, 
Tachygoninae, Thecesterninae, Trypetidinae: C. W. O’Brien, 1997 
Curculionidae: Curculioninae: L. Gibson, 1985 
Curculionidae: Erirhininae: C. W. O’Brien and D. M. Anderson, 1996 
Curculionidae: Hylobiinae: C. W. O’Brien, 1986 
Curculionidae: Pissodinae: L. F. O’Brien, 1989 
Curculionidae: Polydrosinae: Tanymecini: A. T. Howden, 1993 
Derodontidae: J. F. Lawrence, 1989 
Endomychidae: H. F. Strohecker, 1986 
Heteroceridae: F. Pacheco, 1978 
Ithyceridae: C. W. O’Brien, 1994 
Languriidae: J. F. Lawrence & P. Vaurie, 1983 
Limnichidae: D. P. Wooldridge, 1986 
Micropeplidae: J. M. Campbell, 1984 
Mordellidae: D. E. Bright, 1986 
Nemonychidae: R. Hamilton, 1994 
Platypodidae: S. L. Wood, 1979 
Pselaphidae: D. Chandler, 1997 
Psephenidae: H. P. Brown, 1983 
Pyrochroidae: D. K. Young, 1982 
Scarabaeidae: Dynastinae and Rutelinae: A. Hardy, 1991 
Scarabaeidae: Geotrupinae: H. F. Howden, 1984 
Silphidae: S. B. Peck and S. E. Miller, 1993 
 
DIPTERA PUBLICATIONS 
 
Atchley, Wirth, Gaskins, and Strauss (compilers). 1981. Bibliography and Keyword Index of the Biting 
Midges (Diptera: Ceratopogonidae), Bibliographies and Literature of Agriculture, No. 13. 
 
Foote, R. H. 1981. Genus Rhagoletis Loew South of the U. S. (Diptera: Tephritidae). U.S.D.A Tech. 
Bulletin 1607. 
 
Foster, G.A. and W. N. Mathis. 2003. A Revision of the Genera Pelomyia Williston and Masoniella 
Vockeroth (Diptera: Tethinidae). Smithsonian Contributions to Zoology, 619:1-63. 
 
Mathis, W. N. and W. W. Wirth. 1977. A New Genus of Psilopine flies (Diptera: Ephydridae) with Notes 
on its Relationships. Proceedings of the Entomological Society of Washington, 79(1): 63-74 [14 figs.]. 
 
Mathis, W. N. 1977. Key to the Neotropical Genera of Parydrinae with a Revision of the Genus Eleleides 
Cresson (Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 90(3): 553-565 [13 
figs.]. 
 
Mathis, W. N and W. W. Wirth. 1977. New Synonymy in Ephydridae (Diptera). Proceedings of the 
Entomological Society of Washington, 79(4): 520. 
 
Mathis, W. N. 1977. A New Synonym and Lectotype Designation in Ephydridae (Diptera). Proceedings 
of the Entomological Society of Washington, 79(4): 537. 
 
Mathis, W. N. 1977. A Revision of the Genus Rhysophora Cresson with a Key to Related Genera 
(Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 90(4): 921-945 [29 figs.; 1 
table]. 
 
Mathis, W. N. 1978. A Revision of the Neartic Species of Limnellia Malloch (Diptera: Ephydridae). 
Proceedings of the Biological Society of Washington, 91(1): 250-293 [50 figs.]. 
 
Mathis, W. N. 1978. New Synonymy in Ephydridae (Diptera). Entomological News, 89(2and3): 105-106. 
 
Mathis, W. N. 1978. A Synopsis of Neotropical Eleleides Cresson (Diptera: Ephydridae). Proceedings of 
the Entomological Society of Washington, 80(4): 465-472 [5 figs.; 1 table]. 
 
Mathis, W. N. and W. W. Wirth. 1978. Paracanace New Genus near Canaceoides Cresson, Three New 
Species, and Notes on their Classification (Diptera: Canacidae). Proceedings of the Entomological Society 
of Washington, 80(4): 524-537 [12 figs.; 1 table]. 
 
Mathis, W. N. 1979. Studies of Notiphilinae (Diptera: Ephydridae), I: Revision of the Neartic Species of 
Notiphila Fallen, Excluding the Caudata Group. Smithsonian Contributions to Zoology, 287: 1-111. 
 
Mathis, W. N. 1979. [Review]. Record of My Life-Work in Entomology, 1903-1904, by C. R. Osten 
Sacken, with an appreciation and introductory preface by K. G. V. Smith. Proceedings of the 
Entomological Society of Washington, 81(2): 330-331. 
 
Mathis, W. N. 1979. Ecuadorian Parascatella (Diptera: Ephydridae). Proceedings of the Entomological 
Society of Washington, 81(4): 602-610 [11 figs.]. 
 
Mathis, W. N. and W. W. Wirth. 1979. Beach flies of Madagascar (Diptera: Canacidae). Annals of the 
Natal Museum, 23(3): 785-796 [16 figs.]. 
 
Mathis, W. N. 1980. Studies of Ephydrinae (Diptera: Ephydridae), III: Revisions of Some Neotropical 
Genera and Species. Smithsonian Contributions to Zoology, 303: 1-50 [77 figs.]. 
 
Mathis, W. N. and G. S. Steyskal. 1980. A Revision of the Genus Oedoparena Curran (Diptera: 
Dryomyzidae: Dryomyzinae). Proceedings of the Entomological Society of Washington, 82(3): 349-359 
[7 figs.]. 
 
Mathis, W. N. and A. Freidberg. 1980. A New Species of Nesopsilopa from Florida (Diptera: 
Ephydridae). Florida Entomologist, 63(3): 279-281 [2 figs.]. 
 
Mathis, W. N. and W. W. Wirth. 1981. Studies of Ephydrinae (Diptera: Ephydridae), IV: Revision of the 
Australian Species of Subgenus Neoscatella Malloch. Smithsonian Contributions to Zoology, 325: 1-27 
[46 figs.]. 
 
Thompson, F. C. and W. N. Mathis. 1981. Haliday's Generic Names of Diptera First Publised in Curtis' 
A Guide to...British Insects (1837). Journal of the Washington Academy of Sciences, 70(2): 80-89. 
 
Mathis, W. N. and K. W. Simpson. 1981. Studies of Ephydrinae (Diptera: Ephydridae), V: Systematics, 
Phylogeny, and Natural History of the Genera Cirrula Cresson and Dimecoenia Cresson in North 
America. Smithsonian Contributions to Zoology, 329: 1-51 [119 figs.]. 
 
Mathis, W. N. and W. W. Wirth. 1981. Hydrellia tritici (Diptera: Ephydridae), New to Hawaii. 
Proceedings of the Hawaiian Entomological Society, 23(3): 371-373. 
 
Mathis, W. N. 1981. Ephydridae. pages 312-316. in: Aquatic Biota of Tropical South America. 
Hurlbert, S. H., (Ed.), University of San Diego Press, San Diego. 
 
Mathis, W. N. 1982. Studies of Ephydrinae (Diptera: Ephydridae), VI: Review of the Tribe Dagini. 
Smithsonian Contributions to Zoology, 345: 1-30. 
 
Mathis, W. N. 1982. Canacidae of Isreal, with a Revison of the Paleartic Species of the Genus Canace 
Haliday (Diptera). Entomologica Scandinavica, 13: 57-66. 
 
Mathis, W. N. 1982. Studies of Ephydrinae (Diptera: Ephydridae), VII: Revision of the Genus Setacera 
Cresson. Smithsonian Contributions to Zoology, 350: 1-57 [138 figs.]. 
 
Mathis, W. N. 1982. Description of a New Species of Nocticanace Malloch (Diptera: Canacidae) from 
Sri Lanka, with Notes on Two Related Species. Proceedings of the Entomological Society of Washington, 
84(3): 421-425. 
 
Mathis, W. N. and A. Freidberg. 1982. New Beach Flies of the Genus Xanthocanace Hendel with a 
Review of the Species from the Western Paleartic (Diptera: Canacidae). Memoirs of the Entomological 
Society of Washington, 10: 97-104. 
 
Mathis, W. N. 1982. Ephydridae. pages 469-472. in: Aquatic Biota of Mexico, Central America and the 
West Indies. Hurlbert, S. H. and Villalobos-Figueroa, A., (Ed.), University of San Diego Press, San 
Diego. 
 
Mathis, W. N. and F. C. Thompson (Eds.). 1982. Recent Advances in Dipteran Systematics - Collections 
of Papers in Honor of C. W. Sabrosky. Memoirs of the Entomological Society of Washington. 
 
Mathis, W. N. 1983. A revision of the genus Dagus Cresson (Diptera: Ephydridae). Proceedings of the 
Entomological Society of Washington, 85(4): 717-726. 
 
Mathis, W. N. 1983. Notes on Brachydeutera Loew (Diptera: Ephydrididae) from North America. 
Entomological News, 94(3): 177-180. 
 
Mathis, W. N. 1984. Notes on the Shore Fly Genus Diedrops (Diptera: Ephydridae). Proceedings of the 
Entomological Society of Washington, 86(2): 349-353 [6 figs.]. 
 
Mathis, W. N. 1984. A Revision of the Genus Asmeringa Becker (Diptera: Ephydridae). Isreal Journal of 
Entomology, 17: 67-79 [21 figs.]. 
 
Mathis, W. N. 1984.A Revision of the Shore Fly Genus Homalometopus Becker (Diptera: Ephydridae). 
Proceedings of the Biological Society of Washington, 97(2): 253-264. 
 
Mathis, W. N. and K. D. Ghorpade. 1985. Studies of Parydrinae (Diptera: Ephydridae), I: A Review of 
the Genus Brachydeutera Loew from the Oriental, Australian, and Oceanian regions. Smithsonian 
Contributions to Zoology, 406: 1-25 [27 figs.]. 
 
Mathis, W. N. 1985. A New Psilopine Genus and Species from Isreal with a Recharacterization of the 
Tribe and Key to Other Psilopine Genera of the Middle East (Diptera: Ephydridae). Proceedings of the 
Entomological Society of Washington, 87(2): 375-380. 
 
Mathis, W. N, 1985. Studies of Parydrinae (Diptera: Ephydridae), II: A Revision of the Shore Fly Genus 
Pelinoides Cresson. Smithsonian Contributions to Zoology, 410: 1-46 [59 figs.]. 
 
Mathis, W. N. and W. E. Steiner. 1985. An Adventive Species of Bracydeutera Loew in North America 
(Diptera: Ephydridae). Journal of the New York Entomological Society, 94(1): 56-61. 
 
Mathis, W. N. 1986. A Revision of the Subgenus Eremotrichoma Soika of the Shore Fly Genus 
Allotrichoma Becker (Diptera: Ephydridae). Isreal Journal of Entomology, 19: 127-139. 
 
Mathis, W. N. 1986. Studies of Psilopinae (Diptera: Ephydridae), I: A Revision of the Shore Fly Genus 
Placopsidella Kertész. Smithsonian Contributions to Zoology, 430: 1-30. 
 
Freidberg, A. and W. N. Mathis, 1986. Studies of Terelliinae (Diptera: Tephritidae): A Revision of the 
Genus Neaspilota Osten Sacken. Smithsonian Contributions to Zoology, 439: 1-75 [204 figs.]. 
 
Mathis, W. N. and C. L. Hogue. 1986. Description of a New Species of the Shore Fly Genus Diedrops 
(Diptera: Ephydridae) from Colombia. Contributions in Science (Los Angeles County Museum), 377: 21-
26. 
 
Wirth, W. W., W. N. Mathis, and J. R. Vockeroth. 1987. 98. Ephydridae. pages 28: 1027-1047. in: 
Manual of the Families and Genera of Neartic Diptera Volume 2. McAlpine, F. M. and Et al, (Eds.), 
[Publ. not given], [Place not given], . 
 
Mathis, W. N. and J. C. Deeming. 1987. A Revision of the Shore Fly Genus Elephantinosoma Becker 
(Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 89(3): 562-570. 
 
Mathis, W. N. and T Zatwarnicki. 1988. Studies on the Shore Fly Tribe Dagini (Diptera: Ephydridae). 
Proceedings of the Entomological Society of Washington, 90(1): 106-121. 
 
Mathis, W. N. 1988. The First Record of the Genus Procanace Hendel from North America, with the 
Description of a New Species (Diptera: Canacidae). Proceedings of the Entomological Society of 
Washington, 90(3): 329-333. 
 
Mathis, W. N. 1988. First Record of the Shore-Fly Genus Placopsidella Kertész from North America 
(Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 90(3): 334-337. 
 
Mathis, W. N. 1988. Beach Flies of the Republic of Seychelles (Diptera: Canacidae). Bulletin of the 
Washington Biological Society, 8: 22-29. 
 
Mathis, W. N. and Z. Y. Jin, Z. Y. 1988. A Review of the Asian Species of the Genus Lamproscatella 
Hendel (Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 101(3): 540-548. 
 
Mathis, W. N. and Z. Y. Jin. 1989. A Review of the Shore Fly Genus Polytrichophora Cresson from 
Asia (Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 102(2): 434-446 [25 
figs.]. 
 
Mathis, W. N. 1989. A Review the Beach Flies of the Carribean and Gulf of Mexico (Diptera: 
Canacidae). Proceedings of the Biological Society of Washington, 102(3): 590-608 [56 figs.]. 
 
Mathis, W. N. and T. Zatwarnicki. 1990. A Revision of the Western Paleartic Species of Athryoglossa 
(Diptera: Ephydridae). Transactions of the American Entomological Society, 116(1): 103-133 [31 figs.]. 
 
Mathis, W. N. 1990. A Revision of the Shore-Fly Genus Diphuia Cresson (Diptera: Ephydridae). 
Proceedings of the Entomological Society of Washington, 92(4): 746-756 [22 figs.]. 
 
Mathis, W. N. 1991. Studies of Gymnomyzinae (Diptera: Ephydridae), II: A Revision of the Shore-Fly 
Subgenus Pseudohecamede Hendel of the Genus Allotrichoma Becker. Smithsonian Contributions to 
Zoology, 522: 1-28 [26 figs.]. 
 
Mathis, W. N. 1992. The First Shore Fly of the Genus Glenanthe Haliday from the Australasian Region 
(Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 94(1): 78-82 [3 figs.]. 
 
Mathis, W. N. and T. Zatwarnicki. 1992. Ephydridae Zetterstedt, 1837 (Insecta, Diptera): Proposed 
Precedence over Gymnomyzidae Latreille, 1829. Bulletin of Zoological Nomenclature, 49(2): 133-136. 
 
Mathis, W. N. 1992. World Catalog of the Beach-Fly Family Canacidae (Diptera). Smithsonian 
Contributions to Zoology, 536: 1-18. 
 
Mathis, W. N. 1993. A Revision of the Shore-Fly Genera Hostis Cresson and Paratissa Coquillett 
(Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 95(1): 21-47 [54 figs.]. 
 
Mathis, W. N. 1993. A New Species and Subgenus of Periscelis Loew from Australia (Diptera: 
Periscelididae). Journal of the Australian Entomological Society, 32: 13-19 [4 figs.]. 
 
Grimaldi, D. A. and W. N. Mathis. 1993. Fossil Periscelididae (Diptera). Proceedings of the 
Entomological Society of Washington, 95(3): 383-403 [17 figs.]. 
 
Mathis, W. N. and T. Zatwarnicki. 1993. A Revision of the Shore-Fly Genus Chlorichaeta Becker 
(Diptera: Ephydridae). Tijdschrift voor Entomologie, 136(1): 55-76 [53 figs.]. 
 
Mathis, W. N. 1994. A Revision of the Shore-Fly Genus Clasiopella Hendel (Diptera: Ephydridae). 
Proceedings of the Entomological Society of Washington, 96(3): 454-465 [25 figs.]. 
 
Baptista, A. and W. N. Mathis. 1994. A Revision of the New World Cyamops Melander (Diptera: 
Periscelididae). Smithsonian Contributions to Zoology, 563: 1-25 [59 figs.]. 
 
Mathis, W. N. 1995. Studies of Gymnomyzinae (Diptera: Ephydridae), VI: A Revision of the Genus 
Glenathe Haliday from the New World. Smithsonian Contributions to Zoology, 567: 1-26 [59 figs.]. 
 
Rao, N. and W. N. Mathis. 1995. A Revision of the Shore-fly Genus Rhysophora Cresson (Diptera: 
Ephydridae). Proceedings of the Entomological Society of Washington, 97(3): 666-688 [35 figs.; 1 table]. 
 
Mathis, W. N. 1995. Shore Flies of the Galápagos Islands (Diptera: Ephydridae). Annals of the 
Entomological Society of America, 88(5): 627-640 [26 figs.; 1 table]. 
 
Mathis, W. N. and T. Zatwarnicki. 1995. World Catalog of Shore Flies (Diptera: Ephydridae). Memoirs 
on Entomology, International, Associated Publishers, 4: 1-423. 
 
Mathis, W. N. and L. Munari. 1996. World Catalog of the Family Tethinidae (Diptera). Smithsonian 
Contributions to Zoology, 584: 1-27 [1 fig.]. 
 
Mathis, W. N. 1997. A Review of the Hoguei Group of Paracanace Mathis and Wirth (Diptera: 
Canacidae). Memoirs of the Entomological Society of Washington, 18: 140-148 [20 figs.]. 
 
Mathis, W. N. 1997. A Review of the Shore-Fly Genus Diphuia Cresson (Diptera: Ephydridae). 
Proceedings of the Entomological Society of Washington, 99(1): 28-36 [17 figs.]. 
 
Mathis, W. N. 1997. A Revision of Neotropical Ditricohphora Cresson (Diptera: Ephydridae). 
Proceedings of the Entomological Society of Washington, 99(4): 697-704 [4 figs.]. 
 
Mathis, W. N. 1997. Shore Flies of the Belizean Cays (Diptera: Ephydridae). Smithsonian Contributions 
to Zoology, 592: 1-77 [258 figs.; 6 tables]. 
 
Mathis, W. N. and T. Zatwarnicki. 1998. A Review of the West Indian Species of Mimapsilopa Cresson 
(Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 100(1): 7-24 [33 figs.]. 
 
Mathis, W. N. and G. A. Foster. 1998. A Revision of the Family Tethinidae (Diptera) From the 
Caribbean, Gulf of Mexico, and Bermuda. Proceedings of the Entomological Society of Washington 
100(4), pp. 601-632 [34 figs] 
 
Mathis, W.N . 1999. A Review Of The Beach-Fly Genus IsocanaceMathis (Diptera: Canacidae). 
Proceedings of the Entomological Society of Washington 101(2),1999, pp. 347-358 [31 figs] 
 
Mathis, W. N, and J. F. Edmiston. 2000. A revision of the Shore-fly genus Lemnaphila Cresson (Diptera: 
Ephydridae). Proceedings of the Entomological Society of Washington 102(3): 652-677 [27 figs.; 3 
tables] 
 
Mathis, W. N. and G. A. Foster. 2000. Notes On Neotropical Species of Tethina Haliday (Diptera: 
Tethinidae). Proceedings of the Entomological Society of Washington 102(3), 2000, pp. 542-548. [4 figs] 
 
Mathis, W. N. and L. Marinoni. 2000. A Cladistic Analysis of Sciomyzidae Fallen (Diptera). Proceedings 
of the Entomological Society of Washington 113(1):162-209. [324 figs] [1 table] 
 
Baptista, A. R. P. and W. N. Mathis. 2000. Notes on the genus Cyamops Melander (Diptera: 
Periscelididae), including description of ten new species. Proceedings of the Entomological Society of 
Washington 102(3): 481-506 [37 figs.] 
 
Mathis, W.N., A. H. Kirk-Spriggs, J. W. Ismay, M. Ackland, J. Rohacek, G. A. Foster, T. Pape, P. S. 
Cranston and R. Meier. 2001. Inter-Tidal Diptera of Southwestern Africa (Chironomidae, Canacidae, 
Chloropidae, Milichiidae, Tethinidae, Ephydridae, Sphaeroceridae, Coelopidae, Sarcophagidae and 
Anthomyiidae. Cimbebasia 17, pp.85-135 [81 figs] 
 
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